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The crystal in the main room of the Temple is used for a particular ceremony which Loretta Fleice will give you (this is the Arch-Druid quest, I believe). Other than that, it's not all that useful. 

If you have high enough perception you can get a crystal to sell. If you can get to the crystal, then you've done everything you need to do as far as the shrines. Now you have to get the second druid quest from Loretta Fleiss, and visit that last shrine (the one that holds the crystal) at midnight on a full moon. 

The crystal, though, in and of itself, is unimportant.

Also try going to version 1.1, it seems that there was a bug with this part of the game that can stop you from getting to the last crystal. Install the patch and see if that makes a difference.

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